Other than a token mention of web frameworks at the end, this article could have been written 25 years ago. Which is sad, because Perl web development has moved on a lot in that time,
These days, the best way to write a Perl web application is writing a PSGI application, using the Plack toolkit (or something built on top of that). If you do that, you can easily deploy your program as a CGI program, using tools like mod_perl and FastCGI or (my usual choice) as a standalone server behind a proxy server like NGINX.
You say "Perl may not always be the first choice for web development in today’s landscape". Sadly, I think a lot of that is because people still write articles like this which make people think Perl web development is still stuck in the first dotcom boom.